Donatello is one of DIO's illegitimate sons, and the half-brother of Ungalo, Rikiel, and Giorno Giovanna. When he fatefully meets Enrico Pucci, he awakens his Stand, Under World, and battles Jolyne Cujoh and Ermes Costello in Orlando. Although he cooperates with Pucci, he has his own ambitions.
Donatello Versus has short light hair with a black stripe in the middle which extends to the back of his head in the shape of a 'Y'. He has long sideburns which point toward his mouth, blunt bangs that cover his entire forehead, and two ponytails.
Versus wears a choker necklace that has a spiral pattern in the center. He sports a deep V-neck long-sleeved shirt along with matching pants. The front and sides of his shirt and pants have an excessive amount of thick loops. He has shoulder straps going through three front loops on his shirt, as well as two more straps from his belt which travel downward through the front loops on his pants.
Donatello's physical measures, such as height, foot size and blood pressure, are noted to be eerily similar to those of Enrico Pucci.
Beaten down by life, Donatello Versus developed a grudge against society and a desire to become happy at any cost. Having inherited DIO's ruthlessness and selfishness, Donatello lacks his father's manipulative or psychopathic tendencies, and as such even though he is more outwardly well-adjusted, Versus is also much more rash and impulsive.
Donatello isn't opposed to cooperation or even following orders, as at first he put his trust in and tried to mantain a calm, respectful relationship with Enrico Pucci. However, having been pushed around by authority figures and enduring grievous suffering through no fault of his own left Donatello with little tolerance for disrespect and a lot of pride. Feeling like he's being used, he verbally snaps at Pucci when the priest begins doubting Donatello's abilities, and eventually betrays him, considering it his right to inherit Dio's plans in Pucci's place.
Donatello is as ruthless as he is desperate, equally ready to kill children and put his own life on the line to achieve victory. Combined with his single-minded drive and a tendency to loudly express his emotions Donatello can get easily blindsided. When caught off guard Versus will crumble under the pressure, as he fails to kill Jolyne and Ermes after Pucci's repeated comments acknowledging their strength, and eventually gets cornered and forced into submission by them. Donatello is also quite hot-blooded, as he easily blows up on people, gets frustrated quickly and showcases exaggerated facial expressions from smug giddiness to triumphant screams of joys.
- Main article: Under World
Under World allows Donatello to revive past events from Florida's soils and imprison people in them, forcing his victim to experience whatever happened.
Donatello Versus was raised by his mother and stepfather, a man who "did nothing but tell long stories of his womanizing ways," who his mother married at an unknown point in Versus' life.
While running away from home at age 13, a pair of baseball shoes mysteriously fell from the sky and landed on his head. Wondering where they came from, he tried them on. They were way too big, but he decided to keep them anyway because they smelled very good and he liked the clacking noise the spikes made whenever he walked. It wouldn't be long before he was arrested and taken to court, where he learned the shoes belonged to real-life Seattle Mariners outfielder Ichiro Suzuki, originally to be given to disabled children as a gift. Upon hearing the boy say that the shoes fell from the sky, the judge flew into a rage, disgusted by the fact that a 13-year-old boy would stoop so low as to steal from children less fortunate than himself. Versus was quickly deemed guilty on circumstantial evidence of stealing the shoes and was sent to a juvenile detention facility for six months, Versus' mother and stepfather not objecting to his incarceration.
However, four months after Versus' arrest, the true criminal confessed. The criminal was a professional burglar, who, after stealing the shoes, was afraid that he would be caught because the shoes were so famous, and dropped the shoes down a ventilation shaft on top of a building to get rid of them.
When word reached the facility he was being held in that he had been wrongly condemned, Versus had already been physically and emotionally drained, and he was almost too weak to even move because of a string of strange events that had occurred over the course of his incarceration. For instance, Versus tripped while walking in the field of the facility and his hand was impaled by a knife that had been buried there sometime before. Versus would find out the next day that the knife belonged to a corrupt guard who had been hiding it, presumably to threaten other inmates. As a result of this, the guard bullied and beat Versus regularly. His hand wound was finally treated two weeks after the fact, but worms and pus still infested the wound. Possibly as a result of this, Versus fell into a fever and nearly died. Another occurrence was that Versus urinated on a decrepit wall that crumbled and revealed the skeleton of a long-dead woman.
In 2011, Donatello crossed paths with Enrico Pucci after he was shot in the leg and jumped off the sixth story of a building during a botched burglary and was brought to an hospital. He thus met the priest and his half-brothers Ungalo and Rikiel. Pucci subsequently all revealed them their heritage and Stand powers. While Ungalo and Rikiel headed out to confront the heroes, Donatello kept on recovering at the hospital. Donatello recovers and discovers his ability Under World just in time to attack Jolyne and Ermes, who have reached the hospital following Jolyne's intuition.
He instinctively summons a memory of a boy being shot and the stray bullet attracts the attention of the heroes. Donatello and Pucci subsequently take refuge in a cavern under their room. When Jolyne enters it, Under World traps her in the memory of a plane bound to crash. Jolyne partially exits the airplane and tries to call for help. Under World thus tries to attack the string linking Jolyne to Ermes but Jolyne manages to shoot a pen into its hand, driving it away. Nonetheless, Under World summons a memory of Sports Maxx and lures Ermes inside of the cavern, also trapping her in the plane. Pucci keeps commenting the situation, which frustrates Donatello more and more. When Jolyne opens a door to jump out and attack Donatello, he simply traps her in a jet fighter that will also crash. However, Jolyne rejoins the passenger plane. Wanting to advance further, Donatello sneakily summons a memory of Pucci and steals Weather Report's memory DISC. Jolyne contacts a certain Emporio and learns from him that there are two safe seats. To prevent Jolyne and Ermes from using these, Under World tosses sick children on the seats and the plane eventually crash. However, Ermes has covered the children and herself with the two survivors of the crash and their respective duplicate while Jolyne has unravelled herself to hide in Ermes. Stone Free ties up Donatello and hangs it by his feet, preventing him from using his power. Desperate, Donatello commands Under World to fly to Weather Report and give him back his memory. Weather's sealed ability is unleashed and rainbows invade the city.
While Donatello escapes the hole and the hospital, everyone has to contend with a contagious snail invasion. Donatello thus seeks Emporio, expecting to find Jotaro's memory DISC and discover what Pucci's plan is. He unearths a memory of Jolyne calling Emporio and detects him. He easily takes out the boy but Jolyne and Ermes then crash near Versus and Emporio.
Versus summons the Miami Dolphins to push Jolyne away but she hides inside a pipe, rolls towards Versus, and attacks. When Versus summons a memory of the salt to dehydrate Jolyne's body, her arm is dessicated and Versus obtains Jotaro's memory DISC. Nonetheless, Jolyne has touched Emporio who has touched Versus and thus Versus is turned into a snail. The heroes subsequently capture Versus and force him to drive their car. Unable to properly control the car, Donatello crashes in the middle of the fight between Pucci and Weather Report. This allows Pucci to escape and the priest uses an illusion to disguise Donatello as himself. Jolyne kills Donatello with a violent punch to the face, mistaking him for the priest.
Versus appears in JoJo's Bizarre Adventure: All Star Battle as the guide for the support items shop in Story Mode, using his Stand ability to dig up the player's support item of choice at the cost of gold.
- Versus' backstory is either a homage or a direct lift of the backstory of Stanley Yelnats from the novel, (and later feature film) Holes, written by Louis Sachar. Unlike Stanley, however, Versus' life takes a turn for the worse after he is freed.